Education


  • 1982
    Ph.D., Ecology
    UNIVERSITY OF MINNESOTA, Twin Cities Campus
    Dissertation: Breeding Behavior in the Sharp-tailed Grouse

  • 1971
    M.S., Zoology
    UNIVERSITY OF NORTH DAKOTA, Grand Forks ND
    Thesis: Acoustical Communication in the Sharp-tailed Grouse

  • 1962
    B.S., Medicine
    UNIVERSITY OF NORTH DAKOTA, Grand Forks ND

  • 1960
    B.A., Biology and Chemistry
    MACALESTER COLLEGE, St. Paul MN