Education
- Ph.D. in Physiology and Pharmacology
DUKE UNIVERSITY, Durham NC 1981
Thesis: Nonelectrolye Permeability of Lipid Bilayer Membranes.
- M.S. in Zoology
UNIVERSITY OF BRITISH COLUMBIA, Vancouver BC 1977
Thesis: Total Body Water and Partitioning of Salt Excretion in Glaucous-winged Gulls, Larus glaucesens.
- A.B. in Biology with Honors
GRINNELL COLLEGE, Grinnell IA
