Positions Held


  • 2002 – present
    Chair of Biology Department
    ST. OLAF COLLEGE, Northfield MN

  • 1994 – present
    Professor of Biology
    ST. OLAF COLLEGE, Northfield MN

  • 1991 – 1994
    Associate Professor of Physiology and Biophysics
    WRIGHT STATE UNIVERSITY, Dayton OH

  • 1987 – 1991
    Assistant Professor of Physiology and Biophysics
    WRIGHT STATE UNIVERSITY, Dayton OH

  • 1984 – 1987
    Senior Staff Fellow, Laboratory of Mathematical Biology
    NCI, NIH, Bethesda MD

  • 1981 – 1984
    Guest Worker, Staff Fellow
    LKEM, NIH, Bethesda MD